JERUSALEM – Iranian forces based in Syria fired 20 rockets at Israeli front-line military positions in the Golan Heights early Thursday, the Israeli military said, triggering an Israeli reprisal and further escalating heightened tensions between the two bitter enemies in neighboring Syria.

The Israeli military said its Iron Dome rocket defense system intercepted some of the incoming projectiles, while others caused only minimal damage. There were no Israeli casualties.

Lt. Col. Jonathan Conricus, a military spokesman, said Iran’s Al Quds force had fired the rockets at several Israeli bases, though he would not say how Israel had determined the Iranian involvement. The incoming attack set off air raid sirens in the Israeli-controlled Golan.

Israel “views this Iranian attack very severely,” Conricus told reporters. He said Israel had responded, but did not provide details.

“This event is not over,” he added.

Syria’s state media said Syrian air defenses had intercepted “hostile Israeli missiles” early Thursday that were fired over southwestern Damascus.

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Syrian media earlier said the hostilities began with Israeli fire at Syrian positions in the country’s south from across the border. Pro-Syrian media said Syrian missiles then fired at Israeli forces. One TV station, Beirut-based Al-Mayadeen, said at least 50 missiles were fired from Syria at Israeli forces in the occupied Golan Heights. Syrian media said it was the first time in years that Syrians had fired at Israeli forces in Golan.

Israel has been on heightened alert in recent days, anticipating an Iranian attack following Iranian vows to retaliate for what it says are recent Israeli strikes in Syria targeting Iranian outposts.

Late Tuesday, Syrian state media said Israel struck a military outpost near the capital of Damascus. The Britain-based Syrian Observatory for Human Rights said the missiles targeted depots and rocket launchers that likely belonged to Iran’s elite Revolutionary Guard, killing at least 15 people, eight of them Iranians.

Last month, an attack on Syria’s T4 air base in Homs province killed seven Iranian military personnel. On April 30, Israel was said to have struck government outposts in northern Syria, killing more than a dozen pro-government fighters, many of them Iranians.

Israel has neither confirmed nor denied most of the airstrikes. But for months, it has repeatedly said it will not accept a permanent Iranian military presence in Syria.

In February, Israel shot down what it said was an armed Iranian drone that entered Israeli airspace. Israel responded by attacking anti-aircraft positions in Syria, but an Israeli warplane was shot down during the battle.

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Iranian forces moved into Syria after the outbreak of the Syrian civil war in 2011 to back the forces of President Bashar Assad. As that war winds down, and Assad appears to be headed toward victory, Israel fears that Iran, along with tens of thousands of Iranian-backed Shiite militiamen, will carry out attacks against Israel. President Donald Trump’s announcement Tuesday that the U.S. was withdrawing from the 2015 nuclear deal with Iran has triggered uncertainty and threatened to spark more unrest in the Middle East.

Israeli Prime Minister Benjamin Netanyahu traveled to Moscow on Wednesday to meet with President Vladimir Putin and discuss military coordination in Syria.

Russia has also sent forces to Syria to back Assad. But Israel and Russia have maintained close communications to prevent their air forces from coming into conflict.

Together with Putin, Netanyahu toured a parade celebrating the anniversary of the World War II victory over the Nazis and then met the Russian president at the Kremlin for consultations.

After 10 hours together, Netanyahu said he conveyed Israel’s obligation to defend itself against Iranian aggression.

“I think that matters were presented in a direct and forthright manner, and this is important. These matters are very important to Israel’s security at all times and especially at this time,” he said.

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Israel views Iran as its archenemy, citing Iran’s calls for Israel’s destruction, support for militant groups across the region and growing military activity in neighboring Syria. Israel has warned that it will not allow Iran to establish a permanent military presence in Syria.

Israel’s military went on high alert Tuesday and bomb shelters were ordered open in the Israeli-controlled Golan Heights following reports of “irregular activity of Iranian forces in Syria.” After an uneventful night, the military on Wednesday called on residents to return to “full civilian routine,” meaning studies and excursions would continue as usual, although the shelters would remain open.

Amos Gilead, a retired senior Israeli defense official, told a security conference in the coastal town of Herzliya that Iran’s intentions in Syria meant a wider conflagration may only be a matter of time.

“They want to build a second Hezbollah-stan,” he said, referring to the Iranian-backed Lebanese Shiite militant group that last fought a war with Israel in 2006. “They are determined to do it and we are determined to prevent it. It means we are on a collision course.”

Associated Press writers Sarah El Deeb and Zeina Karam in Beirut contributed to this report.

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